1 | <?xml version='1.0' encoding='UTF-8'?> |
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2 | <NineML xmlns="http://nineml.incf.org/9ML/0.3" |
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3 | xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" |
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4 | xsi:schemaLocation="http://nineml.incf.org/9ML/0.3 ../NineML_v0.3.xsd" |
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5 | name="Brunel (2000) Model A Network with alpha synapses"> |
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6 | |
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7 | <Property><!-- Scales the size of the network (total 5*order neurons) --> |
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8 | <label>order</label> |
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9 | <value>2500</value> |
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10 | </Property> |
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11 | |
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12 | <Property><!-- Number of excitatory neurons --> |
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13 | <label>Ne</label> |
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14 | <value><MathInline>4 * order</MathInline></value> |
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15 | </Property> |
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16 | |
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17 | <Property><!-- Number of inhibitory neurons --> |
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18 | <label>Ni</label> |
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19 | <value><MathInline>1 * order</MathInline></value> |
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20 | </Property> |
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21 | |
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22 | <Property><!-- Connectivity probability --> |
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23 | <label>epsilon</label> |
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24 | <value>0.1</value> |
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25 | </Property> |
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26 | |
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27 | <Property><!-- Total number of excitatory synapses --> |
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28 | <label>Ce</label> |
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29 | <value><MathInline>epsilon * Ne</MathInline></value> |
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30 | </Property> |
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31 | |
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32 | <Property><!-- Total number of inhibitory synapses --> |
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33 | <label>Ci</label> |
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34 | <value><MathInline>epsilon * Ni</MathInline></value> |
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35 | </Property> |
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36 | |
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37 | <Property><!-- Total number of external synapses --> |
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38 | <label>Cext</label> |
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39 | <value><MathInline>Ce</MathInline></value> |
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40 | </Property> |
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41 | |
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42 | <Property><!-- Global delay for all neurons in the group --> |
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43 | <label>delay</label> |
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44 | <value>1.5</value> |
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45 | <unit>ms</unit> |
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46 | </Property> |
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47 | |
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48 | <Property> <!-- Synaptic weight --> |
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49 | <label>J</label> |
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50 | <value>0.1</value> |
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51 | <unit>mV</unit> |
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52 | </Property> |
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53 | |
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54 | <Property> |
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55 | <label>J_max_unit</label> |
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56 | <value><MathInline>1.209</MathInline></value> |
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57 | </Property> |
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58 | |
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59 | <Property> |
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60 | <label>g</label> <!-- Relative strength of inhibitory synapses --> |
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61 | <value>3.0</value> |
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62 | </Property> |
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63 | |
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64 | <Property> <!-- nu_ext / nu_thr --> |
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65 | <label>eta</label> |
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66 | <value>2.0</value> |
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67 | </Property> |
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68 | |
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69 | <Property> <!-- Excitatory weights --> |
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70 | <label>Je</label> |
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71 | <value><MathInline>J / J_max_unit</MathInline></value> |
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72 | </Property> |
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73 | |
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74 | <Property> <!-- Inhibitory weights --> |
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75 | <label>Ji</label> |
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76 | <value><MathInline>-g * Je</MathInline></value> |
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77 | </Property> |
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78 | |
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79 | <Property> <!-- External weights --> |
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80 | <label>Jext</label> |
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81 | <value><MathInline>Je</MathInline></value> |
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82 | </Property> |
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83 | |
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84 | <Property> <!-- Firing threshold --> |
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85 | <label>theta</label> |
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86 | <value>20.0</value> |
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87 | <unit>mV</unit> |
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88 | </Property> |
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89 | |
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90 | <Property><!-- Membrane time constant --> |
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91 | <label>tau</label> |
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92 | <value>20.0</value> |
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93 | <unit>ms</unit> |
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94 | </Property> |
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95 | |
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96 | <Property><!-- Synapse time constant --> |
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97 | <label>tau_syn</label> |
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98 | <value>0.5</value> |
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99 | </Property> |
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100 | |
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101 | <Property> <!-- Threshold rate --> |
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102 | <label>nu_thresh</label> |
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103 | <value><MathInline>theta / (Je * Ce * tau * exp (1.0) * tau_syn)</MathInline></value> |
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104 | </Property> |
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105 | |
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106 | <Property><!-- External rate per synapse --> |
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107 | <label>nu_ext</label> |
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108 | <value><MathInline>eta * nu_thresh</MathInline></value> |
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109 | </Property> |
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110 | |
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111 | <Property> <!-- Mean input spiking rate --> |
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112 | <label>input_rate</label> |
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113 | <value><MathInline>1000.0 * nu_ext</MathInline></value> |
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114 | <unit>Hz</unit> |
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115 | </Property> |
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116 | |
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117 | <Component name="E"> |
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118 | |
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119 | <Definition> |
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120 | <link> BrunelIaF.xml </link> |
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121 | </Definition> |
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122 | |
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123 | <properties> |
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124 | |
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125 | <Property> |
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126 | <label>tau</label> |
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127 | <value>20.0</value> |
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128 | </Property> |
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129 | |
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130 | <Property> |
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131 | <label>theta</label> |
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132 | <value><MathInline>theta</MathInline></value> |
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133 | </Property> |
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134 | |
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135 | <Property> |
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136 | <label>tau_rp</label> |
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137 | <value>2.0</value> |
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138 | </Property> |
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139 | |
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140 | <Property> |
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141 | <label>Vreset</label> |
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142 | <value>10.0</value> |
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143 | </Property> |
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144 | |
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145 | <Property> |
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146 | <label>R</label> |
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147 | <value>1.0</value> |
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148 | </Property> |
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149 | |
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150 | <Initial> |
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151 | <label>Isyn</label> |
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152 | <value>0.0</value> |
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153 | </Initial> |
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154 | |
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155 | <Initial> |
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156 | <label>V</label> |
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157 | <value><MathInline>randomUniform ()</MathInline></value> |
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158 | </Initial> |
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159 | |
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160 | <Initial> |
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161 | <label>t_rpend</label> |
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162 | <value>0.0</value> |
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163 | </Initial> |
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164 | |
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165 | <Initial> |
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166 | <label>spikeOutput</label> |
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167 | <value>false</value> |
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168 | </Initial> |
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169 | |
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170 | <Initial> |
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171 | <label>refractoryEnd</label> |
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172 | <value>false</value> |
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173 | </Initial> |
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174 | |
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175 | </properties> |
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176 | |
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177 | </Component> |
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178 | |
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179 | <Component name="I"> |
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180 | |
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181 | <Definition> |
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182 | <link> BrunelIaF.xml </link> |
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183 | </Definition> |
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184 | |
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185 | <properties> |
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186 | |
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187 | <Property> |
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188 | <label>tau</label> |
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189 | <value>20.0</value> |
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190 | </Property> |
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191 | |
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192 | <Property> |
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193 | <label>theta</label> |
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194 | <value><MathInline>theta</MathInline></value> |
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195 | </Property> |
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196 | |
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197 | <Property> |
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198 | <label>tau_rp</label> |
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199 | <value>2.0</value> |
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200 | </Property> |
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201 | |
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202 | <Property> |
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203 | <label>Vreset</label> |
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204 | <value>10.0</value> |
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205 | </Property> |
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206 | |
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207 | <Property> |
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208 | <label>R</label> |
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209 | <value>1.0</value> |
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210 | </Property> |
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211 | |
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212 | <Initial> |
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213 | <label>Isyn</label> |
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214 | <value>0.0</value> |
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215 | </Initial> |
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216 | |
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217 | <Initial> |
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218 | <label>V</label> |
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219 | <value><MathInline>randomUniform ()</MathInline></value> |
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220 | </Initial> |
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221 | |
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222 | <Initial> |
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223 | <label>t_rpend</label> |
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224 | <value>0.0</value> |
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225 | </Initial> |
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226 | |
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227 | <Initial> |
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228 | <label>spikeOutput</label> |
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229 | <value>false</value> |
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230 | </Initial> |
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231 | |
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232 | <Initial> |
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233 | <label>refractoryEnd</label> |
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234 | <value>false</value> |
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235 | </Initial> |
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236 | |
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237 | |
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238 | </properties> |
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239 | |
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240 | </Component> |
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241 | |
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242 | |
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243 | <Component name="Ext"> |
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244 | |
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245 | <Definition> |
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246 | <link> http://www.NineML.org/stim/Poisson.9ml </link> |
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247 | </Definition> |
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248 | |
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249 | <properties> |
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250 | |
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251 | <Property> |
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252 | <label>rate</label> |
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253 | <value><MathInline>input_rate</MathInline></value> |
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254 | </Property> |
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255 | |
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256 | </properties> |
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257 | |
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258 | </Component> |
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259 | |
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260 | <Component name="ISyn"> |
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261 | |
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262 | <Definition> |
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263 | <link> AlphaPSR.xml </link> |
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264 | </Definition> |
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265 | |
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266 | <properties> |
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267 | |
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268 | <Property> |
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269 | <label>tau_syn</label> |
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270 | <value><MathInline>tau_syn</MathInline></value> |
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271 | </Property> |
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272 | |
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273 | <Initial> |
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274 | <label>A</label> |
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275 | <value>0.0</value> |
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276 | </Initial> |
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277 | |
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278 | <Initial> |
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279 | <label>B</label> |
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280 | <value>0.0</value> |
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281 | </Initial> |
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282 | |
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283 | <Initial> |
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284 | <label>spike</label> |
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285 | <value>false</value> |
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286 | </Initial> |
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287 | |
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288 | <Initial> |
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289 | <label>tspike</label> |
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290 | <value>0.0</value> |
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291 | </Initial> |
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292 | |
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293 | <Initial> |
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294 | <label>Ispike</label> |
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295 | <value>0.0</value> |
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296 | </Initial> |
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297 | |
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298 | <Initial> |
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299 | <label>Isyn</label> |
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300 | <value>0.0</value> |
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301 | </Initial> |
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302 | |
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303 | </properties> |
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304 | |
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305 | </Component> |
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306 | |
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307 | <Component name="ESyn"> |
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308 | |
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309 | <Definition> |
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310 | <link> AlphaPSR.xml </link> |
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311 | </Definition> |
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312 | |
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313 | <properties> |
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314 | |
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315 | <Property> |
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316 | <label>tau_syn</label> |
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317 | <value><MathInline>tau_syn</MathInline></value> |
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318 | </Property> |
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319 | |
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320 | <Initial> |
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321 | <label>A</label> |
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322 | <value>0.0</value> |
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323 | </Initial> |
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324 | |
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325 | <Initial> |
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326 | <label>B</label> |
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327 | <value>0.0</value> |
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328 | </Initial> |
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329 | |
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330 | <Initial> |
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331 | <label>spike</label> |
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332 | <value>false</value> |
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333 | </Initial> |
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334 | |
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335 | <Initial> |
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336 | <label>tspike</label> |
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337 | <value>0.0</value> |
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338 | </Initial> |
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339 | |
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340 | <Initial> |
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341 | <label>Ispike</label> |
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342 | <value>0.0</value> |
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343 | </Initial> |
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344 | |
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345 | <Initial> |
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346 | <label>Isyn</label> |
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347 | <value>0.0</value> |
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348 | </Initial> |
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349 | |
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350 | </properties> |
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351 | |
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352 | </Component> |
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353 | |
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354 | |
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355 | <Component name="ExternalPlasticity"> |
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356 | |
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357 | <Definition> |
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358 | <link> ConstantPlasticity.xml </link> |
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359 | </Definition> |
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360 | |
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361 | <properties> |
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362 | |
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363 | <Initial> |
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364 | <label>weight</label> |
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365 | <value><MathInline>Jext</MathInline></value> |
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366 | </Initial> |
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367 | |
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368 | </properties> |
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369 | |
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370 | </Component> |
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371 | |
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372 | <Component name="ExcitatoryPlasticity"> |
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373 | |
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374 | <Definition> |
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375 | <link> ConstantPlasticity.xml </link> |
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376 | </Definition> |
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377 | |
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378 | <properties> |
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379 | |
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380 | <Initial> |
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381 | <label>weight</label> |
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382 | <value><MathInline>Je</MathInline></value> |
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383 | </Initial> |
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384 | |
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385 | </properties> |
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386 | |
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387 | </Component> |
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388 | |
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389 | <Component name="InhibitoryPlasticity"> |
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390 | |
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391 | <Definition> |
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392 | <link> ConstantPlasticity.xml </link> |
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393 | </Definition> |
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394 | |
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395 | <properties> |
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396 | |
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397 | <Initial> |
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398 | <label>weight</label> |
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399 | <value><MathInline>Ji</MathInline></value> |
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400 | </Initial> |
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401 | |
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402 | </properties> |
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403 | |
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404 | </Component> |
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405 | |
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406 | |
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407 | <Component name="RandomUniform"> |
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408 | |
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409 | <Definition> |
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410 | <link> RandomUniformConnection.xml </link> |
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411 | </Definition> |
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412 | |
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413 | <properties> |
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414 | <Property> |
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415 | <label>epsilon</label> |
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416 | <value><MathInline>epsilon</MathInline></value> |
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417 | </Property> |
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418 | </properties> |
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419 | |
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420 | </Component> |
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421 | |
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422 | <Group name="BrunelCaseAalpha"> |
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423 | |
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424 | <Population name="Exc"> <!-- Excitatory population definition --> |
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425 | <prototype><reference>E</reference></prototype> |
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426 | <number><MathInline>Ne</MathInline></number> |
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427 | </Population> |
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428 | |
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429 | <Population name="Inh"> <!-- Inhibitory population definition --> |
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430 | <prototype><reference>I</reference></prototype> |
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431 | <number><MathInline>Ni</MathInline></number> |
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432 | </Population> |
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433 | |
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434 | <Population name="Ext"> <!-- External sources definition --> |
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435 | <prototype><reference>Ext</reference></prototype> |
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436 | <number><MathInline>Cext</MathInline></number> |
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437 | </Population> |
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438 | |
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439 | <Set name="All neurons"> <!-- The set of all neurons --> |
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440 | <union> |
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441 | <left><singleton>Exc</singleton></left> |
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442 | <right><singleton>Inh</singleton></right> |
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443 | </union> |
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444 | </Set> |
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445 | |
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446 | <!-- Projection from external sources to all neurons --> |
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447 | <Projection name="External" |
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448 | input_event_port="spike" |
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449 | input_analog_port="Isyn" |
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450 | output_event_port="spikeOutput" |
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451 | input_response_port="Ispike"> |
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452 | <source><reference>Ext</reference></source> |
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453 | <target><reference>All neurons</reference></target> |
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454 | <rule operator="all-to-all"/> |
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455 | <response><reference>ESyn</reference></response> |
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456 | <plasticity><reference>ExternalPlasticity</reference></plasticity> |
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457 | </Projection> |
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458 | |
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459 | <!-- Projection from excitatory neurons to all neurons --> |
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460 | <Projection name="Excitation" |
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461 | input_event_port="spike" |
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462 | input_analog_port="Isyn" |
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463 | output_event_port="spikeOutput" |
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464 | input_response_port="Ispike"> |
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465 | <source><reference>Exc</reference></source> |
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466 | <target><reference>All neurons</reference></target> |
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467 | <rule operator="for-each"> |
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468 | <reference>RandomUniform</reference> |
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469 | <state>connection</state> |
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470 | </rule> |
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471 | <response><reference>ESyn</reference></response> |
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472 | <plasticity><reference>ExcitatoryPlasticity</reference></plasticity> |
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473 | </Projection> |
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474 | |
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475 | <!-- Projection from inhibitory neurons to all neurons --> |
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476 | <Projection name="Inhibition" |
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477 | input_event_port="spike" |
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478 | input_analog_port="Isyn" |
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479 | output_event_port="spikeOutput" |
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480 | input_response_port="Ispike"> |
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481 | <source><reference>Inh</reference></source> |
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482 | <target><reference>All neurons</reference></target> |
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483 | <rule operator="for-each"> |
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484 | <reference>RandomUniform</reference> |
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485 | <state>connection</state> |
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486 | </rule> |
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487 | <response><reference>ISyn</reference></response> |
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488 | <plasticity><reference>InhibitoryPlasticity</reference></plasticity> |
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489 | </Projection> |
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490 | |
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491 | <!-- Record spikes from the given set --> |
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492 | <SpikeRecording set="All neurons"/> |
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493 | |
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494 | </Group> |
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495 | |
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496 | </NineML> |
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